Trichoptera Taxonomy Wolfgang Tobias Andreas Allspach Eva-Maria Gerstner Claus Weiland Lothar Menner A working list of the taxon names of the order Trichoptera (caddisflies; Insecta). Animalia Arthropoda Mandibulata Insecta Trichoptera Limnephilidae Molannidae Psychomyiidae Leptoceridae Rhyacophilidae Beraeidae Philopotamidae Brachycentridae Glossosomatidae Hydroptilidae Phryganeidae Polycentropodidae Hydropsychidae Sericostomatidae Lepidostomatidae Ecnomidae Goeridae Thremmatidae Arctopsychidae Limnocentropidae Helicopsychidae Odontoceridae Apatania Pseudopsilopteryx Acrophylax Limnephilus Apatidea Chaetopteryx Stenophylax Hydatophylax Neophylax Allogamus Halesus Psychoglypha Antipodoecia Arctopora Drusus Annitella Asynarchus Micropterna Plectrotarsus Grammotaulius Ulmerochorema Rhadicoleptus Anabolia Chilostigma Brachypsyche Pycnopsyche Potamophylax Pseudostenophylax Metanoea Enoicyla Oncosmoecus Nemotaulius Psilopteryx Amphipsyche Lenarchus Caenota Philanisus Parachiona Glyphotaelius Archaeophylax Conoesucus Melampophylax Cryptothrix Frenesia Ecclisopteryx Oligoplectrum Chaetopterygopsis Homilia Rhyacophila Psilopsyche Synafophora Colpotaulius Mesophylax Xiphocentron Alloecella Agapetus Oxyethira Platyphylax Aethaloptera Caborius Atriplectides Ironoquia Consorophylax Anomalopteryx Phacopteryx Phryganea Potamorites Parastenopsyche Taskiria Stenopsychodes Tasmathrus Eosericostoma Phylloicus Molannodes Molanna Tinodes Paduniella Lype Psychomyia Metalype Asmicridea Athripsodes Oecetis Leptocerus Setodes Triaenodes Mystacides Ceraclea Neuronia Adicella Radema Ymyia Pycnopsyche Neophylax Bimus Ironoquia Thamastes Ylodes Bansiola Notidobia Erotesis Ganonema Rhyacophila Pararhyacophila Ernodes Beraea Beraeodes Beraeamyia Nyctiophylax Philopotamus Wormaldia Phylocentropus Chimarra Dolophilodes Micrasema Brachycentrus Psilopteryx Oligoplectrum Glossosoma Agapetus Synagapetus Mystrophora Bachorema Taschorema Hydroptila Oxyethira Orthotrichia Vigarrha Agraylea Stactobiella Dolophilus Allotrichia Oxytrichia Stactobia Ithytrichia Ptilocolepus Leiochiton Hydropsyche Agrypnia Oligotricha Phryganea Oligostomis Trichostegia Hagenella Semblis Cyrnus Holocentropus Polycentropus Plectrocnemia Beraeodes Dipseudopsis Neureclipsis Hydropsyche Cheumatopsyche Diplectrona Hyporhyacophila Sericostoma Oecismus Notidobia Lepidostoma Dinarthrum Crunoecia Lasiocephala Ecnomus Goera Silo Lithax Thremma Arctopsyche Limnocentropus Helicopsyche Odontocerum Apatania zonella Zetterstedt Apatania wallengreni McLachlan Apatania subtilis Martynov Apatania stylata Navas Apatania stigmatella Zetterstedt Apatania scandinavica Svensson & Tjeder Apatania parvula Martynov Apatania muliebris McLachlan Apatania majuscula McLachlan Apatania jokka Tobias Apatania hispida Forsslund Apatania helvetica Schmid Apatania forsslundi Tobias Apatania fimbriata Pictet Apatania eatoniana McLachlan Apatania dalecarlica Forsslund & Tjeder Apatania cimbrica Nielsen Apatania carpathica Schmid Apatania auricula Curtis Pseudopsilopteryx zimmeri McLachlan Acrophylax zerberus Brauer Acrophylax vernalis Dziedzielewicz Limnephilus xanthodes McLachlan Limnephilus vittatus Fabricius Limnephilus subnitidus McLachlan Limnephilus submonilifer Walker Limnephilus subcentralis Brauer Limnephilus stigma Curtis Limnephilus sparsus Curtis Limnephilus sericeus Say Limnephilus rhombicus Linnaeus Limnephilus quadratus Martynov Limnephilus politus McLachlan Limnephilus picturatus McLachlan Limnephilus pantodapus McLachlan Limnephilus nigriceps Zetterstedt Limnephilus nebulosus Kirby Limnephilus moestus Banks Limnephilus minusculus Banks Limnephilus marmoratus Curtis Limnephilus luridus Curtis Limnephilus lunatus Curtis Limnephilus italicus McLachlan Limnephilus indivisus Walker Limnephilus incisus Curtis Limnephilus ignavus McLachlan Limnephilus hirsutus Pictet Limnephilus guadarramicus Schmid Limnephilus griseus Linnaeus Limnephilus graecus Schmid Limnephilus germanus McLachlan Limnephilus fuscinervis Zetterstedt Limnephilus fuscicornis Rambur Rambur Limnephilus flavospinosus Stein Limnephilus flavicornis Fabricius Limnephilus fenestratus Zetterstedt Limnephilus femoratus Zetterstedt Limnephilus femoralis Kirby Limnephilus extricatus McLachlan Limnephilus externus Hagen Limnephilus elegans Curtis Limnephilus eideli Tobias Limnephilus dubius Stephens Limnephilus dispar McLachlan Limnephilus diphyes McLachlan Limnephilus decipiens Kolenati Limnephilus coenosus Curtis Limnephilus centralis Curtis Limnephilus cenadensis Banks Limnephilus borealis Zetterstedt Limnephilus bipunctatus Curtis Limnephilus binotatus Curtis Limnephilus auricula Curtis Limnephilus algosus McLachlan Limnephilus affinis Curtis Apatidea wallengreni McLachlan Apatidea stigmatella Zetterstedt Apatidea auricula Curtis Chaetopteryx villosa Fabricius Chaetopteryx sahlbergi McLachlan Chaetopteryx obscurata McLachlan Chaetopteryx major McLachlan Chaetopteryx maclachlani Stein Chaetopteryx gessneri McLachlan Chaetopteryx fusca Brauer Chaetopteryx clara McLachlan Stenophylax vibex Curtis Stenophylax rotundipennis Brauer Stenophylax permistus McLachlan Stenophylax nigricornis Pictet Stenophylax mitis McLachlan Stenophylax luctuosus Piller & Mitterpacher Stenophylax latipennis Curtis Stenophylax consors McLachlan Hydatophylax variabilis Martynov Hydatophylax infumatus McLachlan Neophylax ussuriensis Martynov Allogamus uncatus Brauer Allogamus stadleri Schmid Allogamus mendax McLachlan Allogamus ligonifer McLachlan Allogamus hilaris McLachlan Allogamus auricula Curtis Allogamus auricollis Pictet Halesus uncatus Brauer Halesus tesselatus Rambur Halesus stadleri Schmid Halesus rubricollis Pictet Halesus radiatus Curtis Halesus mendax McLachlan Halesus melampus McLachlan Halesus ligonifer McLachlan Halesus digitatus Schrank Psychoglypha ulla Milne Psychoglypha alascensis Banks Antipodoecia turneri Mosely Arctopora trimaculata Zetterstedt Drusus trifidus McLachlan Drusus tenellus Klapalek Drusus rectus McLachlan Drusus plicatus Radovanovic Drusus nigrescens Meyer-Duerr Drusus monticola McLachlan Drusus mixtus Pictet Drusus melanchaetes McLachlan Drusus discophorus Radovanovic Drusus discolor Rambur Drusus destitutus McLachlan Drusus chrysotus Rambur Drusus biguttatus Pictet Drusus annulatus Stephens Annitella thuringica Ulmer Annitella obscurata McLachlan Asynarchus thedenii Wallengren Asynarchus lapponicus Zetterstedt Asynarchus iteratus McLachlan Asynarchus contumax McLachlan Asynarchus coenosus Curtis Asynarchus amurensis Martynov Micropterna testacea Gmelin Micropterna sequax McLachlan Micropterna nycterobia McLachlan Micropterna lateralis Stephens Micropterna fissa McLachlan Plectrotarsus tasmanicus Mosely Grammotaulius submaculatus Rambur Grammotaulius signatipennis McLachlan Grammotaulius sibiricus McLachlan Grammotaulius nitidus Müller Grammotaulius nigropunctatus Retzius Grammotaulius atomarius Fabricius Ulmerochorema stigma Ulmer Rhadicoleptus spinifer McLachlan Rhadicoleptus alpestris Kolenati Anabolia sororcula McLachlan Anabolia nervosa Curtis Anabolia laevis Pictet Anabolia furcata Brauer Anabolia concentrica Zetterstedt Anabolia brevipennis Curtis Chilostigma sieboldi McLachlan Brachypsyche sibirica Martynov Pycnopsyche scabripennis Rambur Pycnopsyche lepida Hagen Pycnopsyche guttifer Walker Potamophylax rotundipennis Brauer Potamophylax nigricornis Pictet Potamophylax millenii Klapalek Potamophylax luctuosus Piller & Mitterpacher Potamophylax latipennis Curtis Potamophylax inermis Moretti & Cianficconi 1994 Potamophylax gambaricus Malicky Potamophylax cingulatus Stephens Potamophylax carpathicus Dziedzielewicz Pseudostenophylax rotundipennis Brauer Metanoea rhaetica Schmid Metanoea flavipennis Pictet Enoicyla reichenbachi Kolenati Enoicyla pusilla Burmeister Oncosmoecus quadrinotatus Banks Nemotaulius punctatolineatus Retzius Psilopteryx psorosa Kolenati Psilopteryx curviclavatus Botosaneanu Amphipsyche proluta McLachlan Lenarchus productus Morton Caenota plicata Mosely Philanisus plebeius Walker Parachiona picicornis Pictet Glyphotaelius pellucidus Retzius Archaeophylax ochreus Mosely Conoesucus norelus Mosely Melampophylax nepos McLachlan Melampophylax mucoreus Hagen Melampophylax melampus McLachlan Melampophylax guttatipennis McLachlan Melampophylax austriacus Malicky Cryptothrix nebulicola McLachlan Frenesia missa Milne Ecclisopteryx madida McLachlan Ecclisopteryx guttulata Pictet Ecclisopteryx guttata Pictet Ecclisopteryx dalecarlica Kolenati Oligoplectrum maculatum Fourcroy Chaetopterygopsis maclachlani Stein Homilia leucophaeus Rambur Rhyacophila laevis Pictet Psilopsyche kolbiana Ulmer Synafophora intermedia Klapalek Synafophora dulkejti Martynov Synafophora altaica Martynov Colpotaulius incisus Curtis Mesophylax impunctatus McLachlan Mesophylax aspersus Rambur Xiphocentron haitiensis Banks Alloecella grisea Banks Agapetus fuscipes Curtis Oxyethira frici Klapalek Platyphylax frauenfeldi Brauer Aethaloptera evanescens McLachlan Caborius dubius Stephens Atriplectides dubia Stephens Ironoquia dubia Stephens Consorophylax consors McLachlan Anomalopteryx chauviniana Stein Phacopteryx brevipennis Curtis Phryganea bipunctata Retzius Potamorites biguttatus McLachlan Parastenopsyche bergeri Martynov Taskiria austera Neboiss Stenopsychodes aureoniger Schmid Tasmathrus angustipennis Curtis Eosericostoma aequispina Schmid Phylloicus aculeatus Blanchard Molannodes zelleri McLachlan Molannodes tinctus Zetterstedt Molanna palpata McLachlan Molanna aterrimus Stephens Molanna angustata Curtis Molanna albicans Zetterstedt Tinodes zelleri McLachlan Tinodes waeneri Linnaeus Tinodes unicolor Pictet Tinodes rostocki McLachlan Tinodes pallidulus McLachlan Tinodes maculicornis Pictet Tinodes kimminsi Sykora Tinodes dives Pictet Tinodes assimilis McLachlan Tinodes algirica Vaillant Paduniella vandeli Decamps Lype reducta Hagen Lype phaeopa Stephens Psychomyia pusilla Fabricius Fabricius Psychomyia flavida Hagen Metalype fragilis Pictet Asmicridea edwardsii McLachlan 1866 Athripsodes wetzeli Ross Athripsodes viridis Fourcroy Athripsodes nigronervosa Retzius Athripsodes fulva Rambur Athripsodes dissimilis Stephens Athripsodes commutatus Rostock Athripsodes cinereus Curtis Athripsodes bilineatus Linnaeus Athripsodes aterrimus Stephens Athripsodes annulicornis Stephens Athripsodes angustipennis Curtis Athripsodes albifrons Linnaeus Oecetis tripunctata Fabricius Oecetis testacea Curtis Oecetis ochracea Curtis Oecetis notata Rambur Oecetis lacustris Pictet Oecetis inconspicua Walker Oecetis furva Rambur Oecetis flavicornis Pictet Oecetis cinerascens Hagen Leptocerus tineiformis Curtis Leptocerus senilis Burmeister 1839 Leptocerus riparius Albarda Leptocerus nigronervosus Retzius Leptocerus lusitanicus McLachlan Leptocerus interruptus Fabricius Leptocerus excisus Morton Leptocerus dissimilis Stephens Leptocerus commutatus Rostock Leptocerus cinereus Curtis Leptocerus bilineatus Linnaeus Leptocerus aterrimus Stephens Leptocerus annulicornis Stephens Leptocerus alboguttatus Hagen Leptocerus albifrons Linnaeus Setodes tineiformis Curtis Setodes punctatus Fabricius Setodes argentipunctellus McLachlan Triaenodes simulans Tjeder Triaenodes reuteri McLachlan Triaenodes internus McLachlan Triaenodes forsslundi Tjeder Triaenodes djaneti Vaillant Triaenodes detruncatus Martynov Triaenodes conspersus Rambur Triaenodes bicolor Curtis Mystacides sibiricus Martynov Mystacides sepulchralis Walker Mystacides rhombicus Linnaeus Mystacides obsoleta Hagen Mystacides nigra Linnaeus Mystacides longicornis Linnaeus Mystacides bifidus Martynov Mystacides azurea Linnaeus Ceraclea senilis Burmeister Ceraclea perplexa McLachlan Ceraclea nigronervosa Retzius Ceraclea fulva Rambur Ceraclea excisa Morton Ceraclea dissimilis Stephens Ceraclea annulicornis Stephens Ceraclea annulatus Stephens Ceraclea alboguttata Hagen Ceraclea albimacula Rambur Neuronia ruficrus Scopoli Adicella reducta McLachlan Adicella filicornis Pictet Radema ovalis Martynov Radema foliata Martynov Radema bellicosa Martynov Ymyia interrupta Fabricius Pycnopsyche guttifer Walker Neophylax fuscus Banks Bimus flavidus McLachlan Ironoquia dubia Stephens Thamastes dipterus Hagen Ylodes detruncatus Martynov Bansiola crotchi Walker Notidobia ciliaris Linnaeus Erotesis baltica McLachlan Ganonema americana Walker Rhyacophila vulgaris Pictet Rhyacophila tristis Pictet Rhyacophila torrentium Pictet Rhyacophila stigmatica Kolenati Rhyacophila simulatrix McLachlan Rhyacophila siltalai Döhler Rhyacophila septentrionis McLachlan Rhyacophila rupta McLachlan Rhyacophila retracta Martynov Rhyacophila relicta McLachlan Rhyacophila pubescens Pictet Rhyacophila praemorsa McLachlan Rhyacophila polonica McLachlan Rhyacophila philopotamoides McLachlan Rhyacophila persimilis McLachlan Rhyacophila pascoei McLachlan Rhyacophila occidentalis McLachlan Rhyacophila obliterata McLachlan Rhyacophila nubila Zetterstedt Rhyacophila nevada Schmid Rhyacophila motasi Botosaneanu Rhyacophila mocsaryi Klapalek Rhyacophila minora Banks Rhyacophila meyeri McLachlan Rhyacophila meridionalis Morton Rhyacophila martynovi Mosely Rhyacophila laevis Pictet Rhyacophila intermedia Klapalek Rhyacophila instabilis Curtis Rhyacophila hirticornis McLachlan Rhyacophila hageni McLachlan Rhyacophila glareosa McLachlan Rhyacophila fuscula Walker Rhyacophila fasciata Hagen Rhyacophila fascala Walker Rhyacophila evoluta McLachlan Rhyacophila dorsalis Curtis Rhyacophila denticulata McLachlan Rhyacophila contracta McLachlan Rhyacophila confinium Botosaneanu Rhyacophila centralis Schmid Rhyacophila aurata Brauer Rhyacophila aquitanica McLachlan Rhyacophila angelieri Decamps 1965 Rhyacophila albardana McLachlan Pararhyacophila intermedia McLachlan Pararhyacophila glareosa McLachlan Ernodes vicinus McLachlan Ernodes nigroaurea Mosely Ernodes articularis Pictet Beraea pullata Curtis Beraea maura Curtis Beraea fontana Wiggins Beraea auresi Vaillant Beraeodes minutus Linnaeus Beraeamyia eideli Döhler Nyctiophylax vestitus Hagen Philopotamus variegatus Scopoli Philopotamus montanus Donovan Philopotamus ludificatus McLachlan Philopotamus cingulatus Stephens Wormaldia triangulifera McLachlan Wormaldia subnigra McLachlan Wormaldia pulla McLachlan Wormaldia occipitalis Pictet Wormaldia moesta Banks Wormaldia mediana McLachlan Wormaldia khourmai Schmid Wormaldia echinata Tobias Wormaldia copiosa McLachlan Phylocentropus placidus Banks Chimarra marginata Linnaeus Dolophilodes distinctus Walker Micrasema vestitum Navas Micrasema tristellum McLachlan Micrasema setiferum Pictet Micrasema scissum McLachlan Micrasema minimum McLachlan Micrasema longulum McLachlan Micrasema gelidum McLachlan Micrasema difficile Mosely Brachycentrus subnubilus Curtis Brachycentrus montanus Klapalek Brachycentrus maculatus Fourcroy Psilopteryx psorosa Kolenati Oligoplectrum maculatum Fourcroy Glossosoma vernale Pictet Glossosoma ussuricum Martynov Glossosoma nylanderi McLachlan Glossosoma nigrior Banks Glossosoma intermedium Klapalek Glossosoma conformis Neboiss Glossosoma boltoni Curtis Agapetus rectigonopoda Botosaneanu Agapetus ochripes Curtis Agapetus numidicus Vaillant Agapetus nimbulus McLachlan Agapetus laniger Pictet Agapetus kirgisorum Martynov Agapetus fuscus Vaillant Agapetus fuscipes Curtis Curtis Agapetus fimbriata Pictet Agapetus delicatulus McLachlan Agapetus comatus Pictet Agapetus bidens McLachlan Synagapetus moselyi Ulmer Synagapetus krawanyi Ulmer Synagapetus iridipennis McLachlan Synagapetus insons McLachlan Synagapetus dubitans McLachlan Synagapetus ater Klapalek Mystrophora intermedia Klapalek Bachorema gisba Mosely Taschorema evansi Mosely Hydroptila vectis Curtis Hydroptila tineoides Dalman Hydroptila tigurina Ris Hydroptila sparsa Curtis Hydroptila simulans Tjeder Hydroptila pulchricornis Pictet Hydroptila occulta Eaton Hydroptila occulata Eaton Hydroptila martini Marshall Hydroptila maclachlani Kimmins Hydroptila lotensis Mosely Hydroptila forcipata Eaton Hydroptila flavicornis Pictet Hydroptila femoralis Kirby Hydroptila dampfi Ulmer Hydroptila angulata Mosely Oxyethira tristella Klapalek Oxyethira tamperensis Malicky Oxyethira simplex Ris Oxyethira sahlbergi McLachlan Oxyethira saggitifera Ris Oxyethira mirabilis Morton Oxyethira klingstedti Nybom Oxyethira frici Klapalek Oxyethira flavicornis Pictet Oxyethira falcata Morton Orthotrichia tragetti Mosely Orthotrichia tetensi Kolbe Orthotrichia costalis Curtis Vigarrha tibialis Navas Agraylea sexmaculata Curtis Agraylea pallidula McLachlan Agraylea multipunctata Curtis Agraylea cognatella McLachlan Stactobiella risi Felber Dolophilus pullus McLachlan Dolophilus copiosus McLachlan Allotrichia pallicornis Eaton Oxytrichia mirabilis Morton Stactobia maculata Vaillant Stactobia maclachlani Kimmins 1949 Stactobia eatoniella McLachlan Stactobia algira Vaillant Ithytrichia lamellaris Eaton Ithytrichia clavata Morton Ptilocolepus granulatus Pictet Leiochiton felina B. Leiochiton fagesii Guinard Hydropsyche contubernalis McLachlan Agrypnia varia Fabricius Agrypnia principalis Martynov Agrypnia picta Kolenati Agrypnia pagetana Curtis Agrypnia obsoleta Hagen Agrypnia czerskyi Martynov Oligotricha striata Linnaeus Oligotricha lapponica Hagen Phryganea striata Linnaeus Phryganea ochrida Malicky Phryganea minor Curtis Phryganea grandis Linnaeus Phryganea bipunctata Retzius Oligostomis reticulata Linnaeus Trichostegia minor Curtis Hagenella clathrata Kolenati Semblis atrata Gmelin Cyrnus trimaculatus Curtis Cyrnus muliebris McLachlan Cyrnus insolutus McLachlan Cyrnus flavidus McLachlan Cyrnus fimbriata Pictet Cyrnus crenaticornis Kolenati Holocentropus stagnalis Albarda Holocentropus picicornis Stephens Holocentropus insignis Martynov Holocentropus dubius Rambur Polycentropus multiguttatus McLachlan Polycentropus irroratus Curtis Polycentropus flavomaculatus Pictet Plectrocnemia geniculata McLachlan Plectrocnemia conspersa Curtis Plectrocnemia conjuncta Martynov Plectrocnemia brevis McLachlan Beraeodes eideli Döhler Dipseudopsis capensis Walker Neureclipsis bimaculata Linnaeus Hydropsyche tobiasi Malicky Hydropsyche tenuis Navas Hydropsyche sparna Ross Hydropsyche smiljae Marinkovic Hydropsyche siltalai Döhler Hydropsyche silfvenii Ulmer Hydropsyche sciligra Malicky Hydropsyche saxonica McLachlan Hydropsyche recurvata Banks Hydropsyche pellucidula Curtis Hydropsyche ornatula McLachlan Hydropsyche orientalis Martynov Hydropsyche newae Kolenati Hydropsyche morosa Ross Hydropsyche morettii de Pietro Hydropsyche modesta Navas Hydropsyche maroccana Navas Hydropsyche instabilis Curtis Hydropsyche incognita Pitsch Hydropsyche guttata Pictet Hydropsyche fulvipes Curtis Hydropsyche fallaciosa Kumanski & Botosaneanu Hydropsyche exocellata Dufour Hydropsyche doehleri Tobias Hydropsyche dissimulata Kumanski & Botosaneanu Hydropsyche dinarica Marinkovic Hydropsyche contubernalis McLachlan Hydropsyche carbonaria McLachlan Hydropsyche bulgaromanorum Malicky Hydropsyche bulbifera McLachlan Hydropsyche botosaneanui Marinkovic Hydropsyche borealis Martynov Hydropsyche angustipennis Curtis Cheumatopsyche sordida Hagen Cheumatopsyche lepida Hagen Cheumatopsyche capitella Martynov Cheumatopsyche antoniensis Malicky Diplectrona ripollensis Tobias Diplectrona felix McLachlan Diplectrona atra McLachlan Hyporhyacophila philopotamoides McLachlan Hyporhyacophila eatoni McLachlan Sericostoma timidum Hagen Sericostoma schneideri Kolenati Sericostoma personatum Kirby & Spence Sericostoma pedemontanum McLachlan Sericostoma galeatum Rambur Sericostoma flavicorne Schneider Oecismus monedula Hagen Notidobia ciliaris Linnaeus Lepidostoma tibialis Carpenter 1933 Lepidostoma prominens Banks Lepidostoma hirtum Fabricius Lepidostoma bryanti Banks Lepidostoma basalis Kolenati Dinarthrum reductum Martynov Crunoecia kempnyi Morton Crunoecia irrorata Curtis Lasiocephala basalis Kolenati Ecnomus tenellus Rambur Goera stylata Ross Goera pilosa Fabricius Silo piceus Brauer Silo pallipes Fabricius Silo nigricornis Pictet Silo mediterraneus McLachlan Lithax obscurus Hagen Lithax niger Hagen Thremma stankovici Marinkovic Thremma gallicum McLachlan Thremma arvernense Giudicelli 1971 Thremma anomalum McLachlan Arctopsyche palpata Martynov Arctopsyche ladogensis Kolenati Arctopsyche callosa Navas Limnocentropus insolitus Ulmer Helicopsyche borealis Hagen Odontocerum albicorne Scopoli